That the key to the present distribution lies in the past had been felt long ago, but at last it was appreciated that the various classes of animals and plants have appeared in successive geological epochs and also at many places remote from each other. The key to the distribution of any group lies in the configuration of land and water of that epoch in which it made its first appearance. Although this sounds like a platitude, it has frequently been ignored. If, for argument's sake, Amphibia were evolved somewhere upon the great southern land-mass of Carboniferous times (supposed by some to have stretched from South America across Africa to Australia), the distribution of this developing class must have proceeded upon lines altogether different from that of the mammals which dated perhaps from lower Triassic times, when the old south continental belt was already broken up. The broad lines of this distribution could never coincide with that of the other, older class, no matter whether the original mammalian centre was in the Afro-Indian, Australian, or Brazilian portion. If all the various groups of animals had come into existence at the same time and at the same place, then it would be possible, with sufficient geological data, to construct a map showing the generalised results applicable to the whole animal kingdom. But the premises are wrong. Whatever regions we may seek to establish applicable to all classes, we are necessarily mixing up several principles, namely geological, historical, i.e. evolutionary, with present day statistical facts. We might as well attempt one compound picture representing a chick's growth into an adult bird and a child's growth into manhood.
In short there are no general regions, not even for each class separately, unless this class be one which is confined to a comparatively short geological period. Most of the great classes have far too long a history and have evolved many successive main groups. Let us take the mammals.
Marsupials live now in Australia and in both Americas, because they already existed in Mesozoic times; Ungulata existed at one time or other all over the world except in Australia, because they are post-Cretaceous;Insectivores, although as old as any Placentalia, are cosmopolitan excepting South America and Australia; Stags and Bears, as examples of comparatively recent Arctogaeans, are found everywhere with the exception of Ethiopia and Australia. Each of these groups teaches a valuable historical lesson, but when these are combined into the establishment of a few mammalian "realms," they mean nothing but statistical majorities. If there is one at all, Australia is such a realm backed against the rest of the world, but as certainly it is not a mammalian creative centre!
Well then, if the idea of generally applicable regions is a mare's nest, as was the search for the Holy Grail, what is the object of the study of geographical distribution? It is nothing less than the history of the evolution of life in space and time in the widest sense. The attempt to account for the present distribution of any group of organisms involves the aid of every branch of science. It bids fair to become a history of the world. It started in a mild, statistical way, restricting itself to the present fauna and flora and to the present configuration of land and water.
Next came Oceanography concerned with the depths of the seas, their currents and temperatures; then enquiries into climatic changes, culminating in irreconcilable astronomical hypotheses as to glacial epochs;theories about changes of the level of the seas, mainly from the point of view of the physicist and astronomer. Then came more and more to the front the importance of the geological record, hand in hand with the palaeontological data and the search for the natural affinities, the genetic system of the organisms. Now and then it almost seems as if the biologists had done their share by supplying the problems and that the physicists and geologists would settle them, but in reality it is not so.
The biologists not only set the problems, they alone can check the offered solutions. The mere fact of palms having flourished in Miocene Spitzbergen led to an hypothetical shifting of the axis of the world rather than to the assumption, by way of explanation, that the palms themselves might have changed their nature. One of the most valuable aids in geological research, often the only means for reconstructing the face of the earth in by-gone periods, is afforded by fossils, but only the morphologist can pronounce as to their trustworthiness as witnesses, because of the danger of mistaking analogous for homologous forms. This difficulty applies equally to living groups, and it is so important that a few instances may not be amiss.
